The animated film tells the background story to the first "Transformers": how friends Orion Pax and D-16 became arch-enemies.
They were once best friends: the Transformers Orion Pax (Optimus Prime) and D-16 (Megatron). They stood up for each other unconditionally on their home planet Cybertron. When the two become embroiled in a conflict with their leader Sentinel Prime, the tide turns: they become bitter enemies who decide the fate of the Transformers in an action-packed showdown ...